https://www.prairiedoghoogland.com/blog-news daily 0.75 2019-01-01 https://www.prairiedoghoogland.com/blog-news/2019/1/1/2otn6jsg1torz474uk4nzg55yicd7g monthly 0.5 2019-01-01 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1546376470723-O28XJ0OMHDU0DLD1R1FU/IMG_E2943.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2019/1/1/om2nbobhcrqymopqhfzn65oql7gkh2 monthly 0.5 2019-01-01 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1546375403128-KJIZGDVRVUM0I8R31X31/VCNP8976.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2019/1/1/4qwa0ar718qfjepxq85ohz3egmg0hj monthly 0.5 2019-01-01 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1546375762251-3M0HC7A7XARS5SM0YOHJ/VCNP8921.JPG Blog & News ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2019/1/1/zp4ixlq014ouuvmjm3tnos8m96fpck monthly 0.5 2019-01-01 https://www.prairiedoghoogland.com/blog-news/2019/1/1/5bid14yqu30m1dji9fmtakt9u86v7n monthly 0.5 2019-01-01 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1546374963842-553GO5LHE9SYISAM9W3U/IMG_1768.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2019/1/1/qc1hl880ljw33wg0gh0l75gi0b60rk monthly 0.5 2019-01-01 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1546374664930-DTIYFJG3X6KK5GEH7SR1/IMG_1324.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2018/4/7/2vdciyevk9hi7z7rbzygpzm93xjeyj monthly 0.5 2018-04-08 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1523103122406-AYNKBO5KEDUPPETR26J2/VCNP0717.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2018/3/25/7rqkvyaamirvge8dxst7qgqfduy49a monthly 0.5 2018-03-25 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1522008170299-QEPSM2XE2UQO9353BA0E/VCNP0126.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2018/3/20/mxo5y4vy6037mf4w8uy9o595st7832 monthly 0.5 2018-03-21 https://www.prairiedoghoogland.com/blog-news/2018/3/18/9q5kfpy5ccpwkbawdcp0l9y21kc2ng monthly 0.5 2018-03-19 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1521418636411-2SR03AVXKQIH2WH4UFXB/VCNP9675.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2018/3/12/745mij4utwq2lxc4aa6x84b6cqocfd monthly 0.5 2018-03-12 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1520864011336-Y1L081Y7OME7RZZQZRRQ/IMG_0995.JPG Blog & News https://www.prairiedoghoogland.com/blog-news/2018/2/28/6a5iynw8hqrho2y5k02hvaxnoxn5z1 monthly 0.5 2018-03-12 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1519836646989-K63LYYWCR4VZ3XXWHBSF/VCNP9404.JPG Blog & News ©MRR 2018 https://www.prairiedoghoogland.com/blog-news/2017/12/14/mfbx48ekxk2ppw5vz1rvnt84324c4i monthly 0.5 2017-12-14 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1513286298349-P28DQO1W6KOC4DG0A8FR/VCNP6010_1.JPG Blog & News - ANNOUNCEMENT https://www.prairiedoghoogland.com/blog-news/2017/11/19/mk5jck3egts5tiy6xaryejrd7bsm33-87gy4 monthly 0.5 2017-12-14 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1511112506484-LM4121SJQUGOVMSZV1EK/IMG_9292.JPG Blog & News - PUBLICATIONS https://www.prairiedoghoogland.com/blog-news/2017/11/2/ubokyrmefnorroelxqmxi3gl9cxjpw monthly 0.5 2018-02-02 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1517587053092-XE76HHX1NPTRTOPE6OUU/IMG_0672.JPG Blog & News - BULLETIN https://www.prairiedoghoogland.com/blog-news/2017/10/25/60ws3wj0h18n5idfszmgqix1q088ho monthly 0.5 2017-10-25 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1508958502323-VYRV0ZIFKDI6O9W6LL6L/IMG_9292.JPG Blog & News - PUBLICATIONS https://www.prairiedoghoogland.com/blog-news/2017/10/12/qrt32opg50ltnpx6gxso0hjwmvlbt7 monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507847439002-7OPLG55C8234CCRCSKYD/IMG_2445_FAV.JPG Blog & News It's alive! ©John Hoogland 2009 https://www.prairiedoghoogland.com/blog-news/2017/9/17/mqb2se6u7avsy6qyazrbch8ivqgtc7 monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505669441270-AHL4RKZQIB5RNS4VRTJ6/VCNP8447.JPG Blog & News Low visibility in Redondo Meadow. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/9/17/5fxeidwdvy4k9mi6uru5q88vp95cjw monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505685240549-RX7OS04L8RYAAJQ3RUQN/VCNP8417.JPG Blog & News A study subject who has lost its markings. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/9/6/oz569cu9vdu3r1eozobtoj1tk85oe7 monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507823495616-ETWQ6P5LM76E4DCZ0A4Z/VCNP8248.JPG Blog & News Head (H) with one of her Hx juveniles. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/9/17/kxi4da8f9df8bx3oeztkg595y2bwom monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505686854742-5TS0YV0MF2ZPCHHOE4X6/VCNP2837_FAV.JPG Blog & News Siblings. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/9/6/pk00eozfrtyg5pq9brvos8cexc9lam monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507823633234-C26N2SYGNVL2W5NALSC8/VCNP8090_1_FAV.JPG Blog & News New post-plague offspring at Redondo Meadow. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/8/31/sg8x7q7ys161u4olklk7undmkwd0y0 monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505686660930-3F7AFD5X30J6BLS54W16/VCNP6086_1_FAV.JPG Blog & News A freshly dyed juvenile. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/8/30/4oyvnst3lzlxlpqjtmf6yu8wgr5xcz monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504139438925-4HU17J17JTBC50RA2JYT/VCNP2600_FAV.JPG Blog & News A baby prairie dog on its first day of emergence aboveground. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/5/12/268jrw8hbrlmcsanjlkuf45iv01k7r monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494637571628-Q6Y9IKCWQ4WJJEP92I4G/image-asset.jpeg Blog & News Squaddies Patrick and Katie read the ultrasound screen as John handles the wand. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/5/12/the-mating-season monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494633621069-XTMT5EKMD6635YSXN19Z/image-asset.jpeg Blog & News Male 2 checks if female Wide Apart is in estrus. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/5/12/2x2gjl754361w7cy8pgdijs9bk6nr1 monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494638503973-MNS2SR5JLEZ317ZVIIDF/image-asset.jpeg Blog & News John and squaddies Patrick and Marlin arrive on site in the morning and collect their gear. ©MRR 2017 https://www.prairiedoghoogland.com/blog-news/2017/1/18/3ss137kmb7rf8hjmkuqdmukdg9wt47 monthly 0.5 2017-10-17 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494638430684-PZD8RZ5DFYKMZOL8LIO0/image-asset.jpeg Blog & News The Valles Grande at the VCNP, New Mexico. ©MRR 2017 https://www.prairiedoghoogland.com/photos-index daily 0.75 2017-10-04 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551165234-SVZ823KLXZJBAE3P7ITG/VCNP1736_FAV.JPG PHOTOS Our first glimpse of emerging prairie dogs is often a head tentatively peeking out from a burrow. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493147391609-8KHTK8EYRWV4CSQLEH6J/VCNP1723.JPG PHOTOS The first prairie dogs to emerge in the spring are typically adult males. Here male 45 surveys his surroundings. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493147826876-GTHTMXAO7EISYL24VWE3/VCNP1751.JPG PHOTOS 18 looks on from his burrow, wary of larger males such as 45, who will spend the spring establishing his territory. Adult males are marked with numbers 0-49. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493147986584-W6OK1A5NRUWPMMREJUHZ/VCNP1888.JPG PHOTOS 18 displays an upright alert posture as he watches for potential threats. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493148103959-1L14G0Q7DR4UEYT4EK2M/VCNP2001.JPG PHOTOS A territorial dispute often involves a chase. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493154117523-YC3O8L2CEG0EGFZKW49I/VCNP2347.JPG PHOTOS Female prairie dogs are marked with numbers 50 and above or with a unique marking, such as four stripes down each flank to designate female 4 Stripe (or 4STR on a datasheet). 4STR was marked in the fall before submerging for the winter, and will lose her marking during her spring molt in May. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493154256877-0NJV1A8B0X8KMO049U2X/VCNP2520.JPG PHOTOS Front paws wet from snow, 45 looks out as he forages. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493154396538-54K569BZTM5DNYC5T3G1/VCNP2848.JPG PHOTOS Female 4 Stripe gathers nesting material for her burrow, likely anticipating a cold night. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493154679696-6SAT46BJ4B8VRTVQ0Y16/VCNP3809.JPG PHOTOS Early in the season (and sometimes well into April), snow cover is still common. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493154756593-LJWM24SEPIAJLH6IIGPQ/VCNP4173.JPG PHOTOS 45 pauses to groom in front of a trap surrounding. Surroundings are set around specific burrows when particular prairie dogs need to be caught for marking in the early Spring. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493154816439-IDUIDTXM2FZNIGJSH2R0/VCNP4326.JPG PHOTOS An unmarked prairie dog has been trapped in a surrounding, ready to be marked before the busy mating season begins. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493154883331-JWPL6XIE8XXIAXFZ2N4R/VCNP4372.JPG PHOTOS Females often stand for long periods at their burrows in the early Spring (Wetsuit here), and will forage near their burrows as the males of their clan run around and often herd them into their burrows to establish dominance. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493155025569-NACF0M06GRMCDUP474K1/VCNP4466.JPG PHOTOS 50 and 83 emerged from the same burrow, possibly sisters, most definitely kin. Prairie dog females exhibit philopatry, remaining in the same clan as their mothers, aunts, and female relatives. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493155476081-70P3RYX29I85N8CFK2AR/VCNP4487.JPG PHOTOS Wetsuit investigates a burrow marking. These tags are placed at heavily-used burrows for referencing when taking behavioral data. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493155539621-9CC7CR8XYCDBDMNZTLTT/VCNP4495.JPG PHOTOS Recently emerged 50 and 83 share a friendly kiss. Context and relationships determine whether kissing is friendly or hostile. After mating, 50 and 83 will become less friendly as they vie for nursery burrows. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493155604611-J172DCEKDASJSH8AOE79/VCNP4621_FAV.JPG PHOTOS 4 Stripe grooms at the mouth of her burrow. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493155698120-GQTE7LQBA4EZI6LCFKQP/VCNP4634.JPG PHOTOS An unmarked dog, having just emerged, will be caught as soon as possible to receive an identification and be entered into the study pool. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493155716465-5AF727ZJM8LSDESAFH7N/VCNP4659.JPG PHOTOS R78, a small yearling male, surveys his surroundings as he emerges from his winter torpor. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505765792569-UHVXAFED9OCR871R3KVJ/1219621.jpg PHOTOS - Early Spring https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505765996661-9JGVLDYJXZW73LTYE3Q8/1219621.jpg PHOTOS - Capture & Handling https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504712791412-S418G62M4X9CHMG929P0/VCNP6136.JPG PHOTOS A closeup of a juvenile's nails during handling. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504712627906-EKJ739VSXXZOYJ79JTSD/VCNP2271.JPG PHOTOS Snow in a trap can inhibit visibility and expose a captured prairie dog to cold and wet. We shake the snow from the traps before opening them in the morning. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725316463-I1FIQ6EXZG8K4UHFZ15P/VCNP2280.JPG PHOTOS John pauses to pose for the camera as he shakes snow from traps. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725116215-K8U34LW3FXHN07MJAYLN/VCNP7388.JPG PHOTOS We use traffic cones to mark important burrows as well as plugging backdoors when attempting to capture a specific prairie dog. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725091001-36HCBIV4NWTQYUTRV4SJ/VCNP7392.JPG PHOTOS A surrounding has been set to capture a prairie dog. When they are proving difficult to catch, we set surroundings just outside the burrow entrance, usually not baited unless we want to provide extra incentive. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725115521-SD1FVRFH0COT3I2FFG55/VCNP7697.JPG PHOTOS Some trap surroundings require up to 36 traps to prevent escape, while smaller ones only require 6. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504887714086-KBSSZ1RRN3OBHFFHW5B3/VCNP4450.JPG PHOTOS The squad works together to relocate an observation blind. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725182634-20XN4VW7W7EEKPA2R13O/VCNP3007.JPG PHOTOS Blinds for the 2017 season can be lifted on and off stilts depending on need for visibility or high winds. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725194701-X947B7TA6LGHVVPZ42KL/VCNP3015.JPG PHOTOS The crew pauses to pose for the camera. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725207820-DLBAYBPNRFONBKKB5MLS/VCNP5322.JPG PHOTOS Squaddie Katie marks the burrow location on flagging tape, which will go on the trap before a newly captured prairie dog is removed and taken to the marking van. Flagging each trap ensures the prairie dog is returned to the correct location. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725270250-AQR59D1L3VQUG9O8LIQG/VCNP2320.JPG PHOTOS Armed with flagging tape, a permanent marker, and a bait bucket, John walks to a surrounding to check the traps. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725249519-6RSO8C0J8P9R0B9GNJTV/VCNP2992.JPG PHOTOS Squaddie Patrick shows Denver Zoo volunteer Tim how to open traps at a surrounding. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725358336-WPL31MCHU5AFZYIV02RJ/IMG_4555.JPG PHOTOS Squaddie Rashidah marks the burrow number on flagging tape before tying the tape to the trap of a captured prairie dog. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725343318-M0GH6I8NP1QGB6AV5PJX/IMG_4570.JPG PHOTOS After labeling the trap with flagging tape, squaddie Rashidah takes a close look at a newly captured prairie dog. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725353401-M2DRV95731SS8INPGW4V/VCNP1683.JPG PHOTOS Arriving on site in the morning, the crew collects their datasheet clipboards and bait buckets before retreating to their blinds/towers for the day. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732512540-BD85GYJGE1NAE4IJPHVV/VCNP0820.JPG PHOTOS Reading the ultrasound machine. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732520046-33CBI6J28E51UE9TWFQJ/VCNP1557.JPG PHOTOS John inspecting traps. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492725347532-JDWX7AE1ZAG7XHAZW8K7/IMG_5767.JPG PHOTOS In the summer, we pull stuffed predators (badgers, coyotes, and bobcats) through the colony to take data on alarm calls. The decoys are pulled using a string at a carefully measured pace. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732522788-VBU47U7R476OWYXJB7H7/VCNP1633.JPG PHOTOS We have dozens upon dozens of traffic cones. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732536618-1D9QOOZQ9YVH7MWUNKGR/VCNP6010_1.JPG PHOTOS Handling is a team effort. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732622470-8QAFFVP9TZTJUZIVJP0H/VCNP6177.JPG PHOTOS Delicately holding a juvenile. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732607693-4ID8KMLLKBNBRJFU6P4U/VCNP6240.JPG PHOTOS Marking a juvenile. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732619867-GN3HVP9X4FU70QEE06Y2/VCNP6336.JPG PHOTOS We use sandwich bags to weigh juveniles. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504825161067-NL6MGFUUARHILPLL2J61/VCNP2857.JPG PHOTOS Squaddie Jimmy engages with a passing family. Speaking with hikers and park visitors gives us the opportunity to explain not only our research but the importance of prairie dogs on the landscape. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505838081128-HYENBNC7J9G94XJP5XMU/IMG_8117.JPG PHOTOS Nursery burrows by mother. Sometimes we have to write notes on our hands for when we're out of our towers. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826467911-4LVTSL4LE5MYDP5H4Q6I/IMG_5227.JPG PHOTOS On busier marking days, captured prairie dogs are placed beneath the marking van to keep them in the cooler shade. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826121785-ICMCSQNX8GGX005S2DD9/IMG_3387.JPG PHOTOS A view of the marking van, captured prairie dogs in wait. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826710739-IGG3JWIUZFRHUO9IA8OM/IMG_5305.JPG PHOTOS Baby prairie dogs wait for handling and marking.©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826891096-VIHHZLUAW7TVWN1L06DN/IMG_4412.JPG PHOTOS A prairie dog remains alert as she waits for handling. ©MRR 2016   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488825849134-GDXPEWV1R3YNRGE8U7ZE/IMG_2390.JPG PHOTOS John checks for eartags before handling. ©MRR 2016   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826251561-EPOSAU4VNTTOTPL17EWM/IMG_3399.JPG PHOTOS A handling bag keeps the prairie dog and handler calm and safe, with a zipper allowing for varying degrees of access to the animal. ©MRR 2016   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826383090-9W6B78FQH6USQX6K78ZW/IMG_3404.JPG PHOTOS The front of the bag can also be opened to allow for head and neck marking. ©MRR 2016   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488825906299-6DM6HNDRTMJKLPMJN5KB/IMG_2394.JPG PHOTOS Squaddie Amber holds the trap as John puts finishing touches to a rear-legs marking. ©MRR 2016   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826818152-QPEBYV9VUA1TGNTZ9ZG6/IMG_4550.JPG PHOTOS Freshly marked prairie dogs are released when the ink dries sufficiently not to smudge. ©MRR 2016   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1488826775963-3OQKD5FQASIDVJQUKLTA/IMG_3241.JPG PHOTOS Some adult females receive unique markings other than a number, such as Wetsuit (WS). ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505838894196-A70F1CZJ5R4OB20MSTTQ/VCNP5985_FAV.JPG PHOTOS Taking a tissue sample. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505838919977-WT689BKKGB3T60UBG99E/VCNP6124_FAV.JPG PHOTOS Weighing a juvenile. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505838946787-1QK4BLNT39AE88IP96V9/VCNP6005_1_FAV.JPG PHOTOS A closeup. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505920802116-VCRRLYAVN319V8SCVKMY/IMG_8355.JPG PHOTOS An unmarked juvenile. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505920773530-V2JCWTPK5XZ90IJ31S7X/IMG_8389_FAV.JPG PHOTOS Returning a marked juvenile. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505920670454-E5R2Y8AQEQ3Q2F8RUJPO/IMG_5844.JPG PHOTOS Eye contact in the trap. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505765952917-F1VKV1FXEO8K0HBF82VR/1219621.jpg PHOTOS - juveniles https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828751825-IC0E5VL8DHL4L1SNZB7W/IMG_2013_FAV.JPG PHOTOS A closeup portrait. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828204755-AHLHSPP2CCK0KIRBQC4Y/IMG_1504.JPG PHOTOS A mother carefully carries her baby in her mouth as she transfers her litter from one burrow into another. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828451491-00C97X9EP6GF8AGDUSOE/IMG_1575.JPG PHOTOS After a little over five weeks nursing underground, this litter makes their first emergence into the sunlight.©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828460218-G3C7JGOKKGASPCJFZF55/IMG_1589.JPG PHOTOS We mark emergent juveniles according to their mother's marking scheme. The following year these individuals will receive unique markings, but for now the only distinguishing mark is a cap of ink on the head for female babies.©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493829675036-TR332HIKSUNBAARLBZ8D/IMG_5403.JPG PHOTOS In a trap. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828508795-HAFC6HFAGDVJRCH3KXWQ/IMG_1596.JPG PHOTOS It's a tough life for a prairie dog, and juveniles are especially vulnerable to predation. Here a long-tailed weasel approaches a nursery burrow.©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828750352-VR2F9PKEBXR36RP0CK2E/IMG_2014.JPG PHOTOS Juvenile prairie dogs are very small.©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828811571-013TR3L9HXF01KO699DQ/IMG_2018.JPG PHOTOS An overhead photo while handling an especially cooperative juvenile.©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828818651-MAFEZSHE3Z5G4C45T4LU/IMG_2100_FAV.JPG PHOTOS When they first emerge, baby prairie dogs will tentatively remain near their burrow.©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828906288-JD5V0WDDS807MRWSDPMK/IMG_2134_FAV.JPG PHOTOS Huddled together. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493828920278-77XPKD4ATC6RZQQGMOIT/IMG_2149_FAV.JPG PHOTOS Babies are tiny but have full bellies after foraging. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493829107602-051VYZINNM2WYBES403M/IMG_4057.JPG PHOTOS Two mothers stand on alert, surrounded by their offspring.©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732034466-LJOKJ85SBHEZCA4JAVH2/IMG_1631_FAV.JPG PHOTOS Mothers may transfer their offspring either by leading them or by carrying them. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504732028673-ZR1GNQIEST0NSUZ5DOAH/IMG_2148_FAV.JPG PHOTOS Relaxed and autogrooming. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809169092-63M8JGSIX84PHT4GEDCR/VCNP2511_FAV.JPG PHOTOS Sibling curiosity. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809207270-HZS84HDPA93E8MYV4O0P/VCNP2868_1_FAV.JPG PHOTOS Testing what's edible. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809198565-DNGNIZKTSLFCJS4V7JQS/VCNP3391_1_FAV.JPG PHOTOS Mother and son both autogrooming (self-grooming). ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809217559-JKNVMIAEZT12IEXDGTQ3/VCNP4241_FAV.JPG PHOTOS A funny face while playing. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809231338-DSTA6AV1Y8WCOACFJJM2/VCNP4605_FAV.JPG PHOTOS Daughter seeks permission to suckle. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809278195-34L6FSVZYSW1WTINEPET/VCNP4778_1_FAV.JPG PHOTOS More common in certain species and less common in others, this Gunnison's prairie dog gives us a rare glimpse of aboveground nursing. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809252808-J58NP6C1ITMOKU44TOTY/VCNP4884_1_FAV.JPG PHOTOS Rearing up to pounce. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809269181-H3JIVGCZRW5951ZLFFXA/VCNP5241_FAV.JPG PHOTOS Tussling in play provides as much learning as it does fun for juveniles. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809296249-BVSO2T66FYHLVPXJZ67K/VCNP5435_FAV.JPG PHOTOS A playful baby pounces on his mother. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809295001-GOMBXZDS77XYCM8NRCEC/VCNP6885_FAV.JPG PHOTOS Who knows what's going on underground; this baby woke up one morning with one eye closed, perhaps from a scratch. Later in the day the eye was open and healthy. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809333752-GN3QFP05GFW93FKBQK1X/VCNP6975_FAV.JPG PHOTOS The Collar Belly marking on this juvenile makes him especially cute - in this researcher's opinion. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809314729-B1IU0QBVR3HKGPO08YGF/VCNP7201_FAV.JPG PHOTOS Siblings tumble into a burrow as they play. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809328043-773VLG6J8P2UXFUAZ4O7/VCNP7233_FAV.JPG PHOTOS Cops and robbers? ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809358867-TIESUX5RLZWX88EOM60I/VCNP7727_FAV.JPG PHOTOS A relaxed sitting juvenile eating grass. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504809351068-8KQHLZOB14RLJOPBUKP8/VCNP7932_FAV.JPG PHOTOS A post-plague juvenile at our Redondo Meadows colony. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810130717-77FJ3RRQXU94SRO27VDS/VCNP2243_1_FAV.JPG PHOTOS A shy litter on their first day of emergence. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810161118-W0UPJ8NJG7ZMEGBFB1JC/VCNP3495_FAV.JPG PHOTOS A baby runs behind his mother during a burrow transfer. This is called "leading." ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810172068-KAE83M6YBZCCQ6V3YRQ3/VCNP4840_FAV.JPG PHOTOS Sibling touch time. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810233472-QWME5EBG3SS2958DE13K/VCNP4873_FAV.JPG PHOTOS A baby drops a glob of dirt after a test bite. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810196468-2QQMVQAQLGGY6D1MYJ0K/VCNP5120_FAV.JPG PHOTOS Released back into his nursery burrow. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810218331-TMOYQ2P9BQV4DA226CO0/VCNP5263_FAV.JPG PHOTOS Sisters playing. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810239069-TW1LFT7I47NYHONSR7TH/VCNP5548_1_FAV.JPG PHOTOS Playing looks nothing like the fights adults will engage it, but play does prepare prairie dogs for hostile interactions in adulthood. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810256963-9KX4I0C1R4Z6ZBHLD4RT/VCNP6574_FAV.JPG PHOTOS A portrait on all fours. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810302652-C4YW4IM5FUGQDCGN0PSU/VCNP6734_FAV.JPG PHOTOS Reaching for the high stuff. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810273409-D9V8T31RUBQ1NB1BUXIR/VCNP7123_FAV.JPG PHOTOS Play copulating is a common form of play among juveniles. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810288667-HTXOEYMRRA9QW8KPWTDX/VCNP7285_FAV.JPG PHOTOS Wrestling. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810304714-XN8VHXYWB1GDXHG5EDAC/VCNP7435_FAV.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810322140-9QROUKX3DI9Q0M10G5AL/VCNP7468_1_FAV.JPG PHOTOS Sometimes three or more babies get in on the play. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810361166-GEMBXI3O8SQQVLMBF1N4/VCNP7480_FAV.JPG PHOTOS Watching juveniles tumble into all sorts of positions can be very amusing. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810344576-J7AJ59H2ESARUZQQZVBT/VCNP7582_1_FAV.JPG PHOTOS A bear hug. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810359470-HQU36ROVD3BRV9FX5KSZ/VCNP7664_FAV.JPG PHOTOS A pounce. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810410237-XCKSV68DRWNZ3FXZDZQ7/VCNP7666_FAV.JPG PHOTOS And a tumble. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810380114-H0ZX3I7CC45ZGUX83BWJ/VCNP6966_FAV.JPG PHOTOS Juveniles spend a lot of time exploring their surroundings, foraging, and playing, but also spend a considerable amount of their day just watching. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810415521-WMJKXMQF7FK1DZWIWTLR/VCNP5307_FAV.JPG PHOTOS Siblings covered in dust as they play-fight over a bit of forage. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810439141-6EC4WI7ELLBDL4I7BCDX/VCNP5105_FAV.JPG PHOTOS Not all juvenile prairie dogs will survive the summer or their first winter. Their carefree early months are brief. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504810396639-XVZ6LAWGZAZL631QQAZ3/VCNP6445_FAV.JPG PHOTOS Two weeks after emerging for the first time, juveniles are already markedly grown. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731059001-3YDC050VPKC3ENZS7O52/IMG_4285_FAV.JPG PHOTOS ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917554578-VOW15EXTO2JP3BVXV0S6/VCNP1862.JPG PHOTOS Males 29 and 45 have a territorial dispute. Many females have emerged in their area, and they will dispute through the mating season, often fighting. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917533162-9IVA4TUCNTPP5WCQRWZ6/VCNP2678.JPG PHOTOS During the mating season, adult males are evidently sexually mature, displaying pigmented testes and a distended scrotum. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917553818-7T1OAZ9PCISS58XFZEMU/VCNP3152_FAV.JPG PHOTOS 45 stands over Wetsuit at her burrow, anticipating her estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917730520-1NG1V0JROJ65ATQRGPPJ/VCNP3191.JPG PHOTOS 45’s pursuit of Wetsuit is relentless on the day she goes into estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918059322-79HWV5RC9Q9VSFYFS9IP/VCNP3750.JPG PHOTOS 45 brings nesting material to a burrow prior to mating with Wetsuit. Though not common, this behavior has been observed in mating males. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918020907-WNGXLBCZWOAWDZNGW1W3/VCNP3777.JPG PHOTOS 45 lets out a mating call at the mouth of the burrow where he has chased Wetsuit. The mating call is sounded before and/or after mating, and if not for context can easily be mistaken for an alarm call. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918626633-Q1NJMLXCGFOPH8E25WQV/VCNP4553.JPG PHOTOS 45 chases and herds 50 into her burrow. We call this a Chase Herd (scored as CH on a datasheet) and it is one of the most common interactions we track, occurring between males, between females, and males to females, though rarely females to males unless that male is a yearling. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918785742-CWHW5COZHTNXM4LLNKYC/VCNP4578.JPG PHOTOS Generally a couple to a few days after mating, females will begin to exhibit maternal behavior, including carrying nesting material to potential nursery burrows. Gestation will last on average 28 days. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918748561-CZ55GUCH8LOPTN5XM7D7/VCNP4600.JPG PHOTOS After mating, 83 licks her vulva. This is one of several aboveground cues that signify a copulation has occurred. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919327047-PP8F7MSUKSD094FV4PV2/VCNP5960.JPG PHOTOS Despite this being her day of estrus, 83 is not ready and rebuffs 45 for now. On their day of estrus, females are only receptive for a few hours. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919486184-S96CE7WO4XNHWVN5B94W/VCNP6300.JPG PHOTOS 24 has a very late wakeup and licks his penis (scored as LP on a datasheet) immediately upon emerging, indicating he spent the morning copulating with a female. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919480313-ZCEMWQW1UOU9WVX0H4MV/VCNP6379.JPG PHOTOS Having already mated with her, 24 tries to keep 6 Stripe underground, chasing and herding her into her burrow to keep her away from other males in the area. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919665906-YX3T8QO020XS5FB9EE7R/VCNP6429.JPG PHOTOS After a male has had his Critical Underground Consortship (CUC) with a female (in which insemination occurs and which is characterized by disappearing underground together for many minutes up to hours), he usually does not go down with her for another copulation. Here 24 and 6 Stripe share a hostile kiss as he tries to keep her at her burrow, having already had their CUC. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919686605-5RH6S55J8U44QUB8S3A6/VCNP6446.JPG PHOTOS 24 will relentlessly chase 6 Stripe to keep her away from other males. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919806245-CYS0VC1CN5RNEAH7DCKS/VCNP8153.JPG PHOTOS R21 chases 49 out of his territory on a day when 5SRS, a female in the area, goes into estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919824286-4U2DSQ43NW9FW4X9IBZ7/VCNP8257.JPG PHOTOS Some time after his first check, 2 comes back for another sniff at Wide Apart. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493920128923-4RKOYK8D09XTIPSCFQET/VCNP9594.JPG PHOTOS Male 36 investigates female 90. Gunnison’s prairie dogs exhibit sexual dimorphism in body mass, and here you can see how much larger 36 is than 90. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493920188437-VS8HVCU3K4QAGQ51F4PZ/VCNP0027_FAV.JPG PHOTOS 36 continues to pursue 90 to check for estrus, and she continues to elude him. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493920201208-ON7IG5WRXV19GBLBDLCD/VCNP0112.JPG PHOTOS RR0, a female who has already mated, brings nesting material to her burrow as she begins to show maternal nesting behaviors. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551737683-82W469V6LK1DDIBRNVSX/VCNP4760.JPG PHOTOS A female runs away from a male trying to sniff her. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551777256-Q6QSL86O0495XI5DC35H/VCNP4615_FAV_LV.JPG PHOTOS Female 4 Stripe licks her vulva on her day of estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551785372-BKR2REUDRHPB8C2KK4XL/VCNP6403.JPG PHOTOS Female 55 runs from male 36 as he pursues her. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551839458-N5RXHIPT173IA2DUISG4/VCNP7882_MC.JPG PHOTOS A mating call from male R21. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551815115-9F0MNLXSFBKQRRO59BY0/VCNP7921_MC.JPG PHOTOS Another mating call from R21. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551833827-LDFXVBXWGX7OFVWE4V4I/VCNP8256.JPG PHOTOS Male 2 comes up behind female Wide Apart to check her for estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551870261-W8Y13KUHQYFY2UWZYVZ1/VCNP8439_FAV_LP.JPG PHOTOS Licking after copulation. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713772960-51SZ8MLRYRTUQ0ZF1OID/IMG_2384_FAV_MC.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713790127-T7KQL76O4KA02L65EGCS/IMG_2375_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713780642-J4N5UNCCDPTPUITGQHPZ/IMG_2438_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731072258-76O9DQYTSZ7UK9IESKTI/IMG_4299_FAV.JPG PHOTOS ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713800949-OV6WF3FDDLRXBK2M3E3P/IMG_2439_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713797881-12PYTA9NVUCT97Q09XAH/IMG_2440_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730766222-KKSAJE2L3WF9SQI5JZ3T/IMG_2603_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730765541-FISPU7RKCP9LCXIPP42T/IMG_2640_FAV_MC.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730793394-7L9173U26PKHJKKZ4QR4/IMG_2644_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730777117-VPPWORPVSA1E7OC1Q0XS/IMG_2647_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730789551-UTIKC0EH9KJVG2TQP2RV/IMG_2696_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730936930-JBNEYMWJRRZNJCRYTOBO/IMG_2707_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730931179-QRZN9MYUMPBVAE00PCL0/IMG_2735_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730951486-4RFIZSINOQJH9LLIXLCB/IMG_3067_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730946815-7EK9U56BEYS7K1AE5WCO/IMG_3324_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730958266-82OCTOEEP5OQFGISJ2JC/IMG_4276_FAV.JPG PHOTOS ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731068539-IJ0BAH29SHA9LTAZUBMD/IMG_4303_1_FAV_MC.JPG PHOTOS ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505765847587-4P72FWY04RO8UIQMUA2H/1219621.jpg PHOTOS - mating behavior https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493932081760-OOIFSZSQVDVQM11TN9AK/VCNP4120.JPG PHOTOS Males 29 and 31 in a hostile kiss. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493932115156-7XOYYRZ7EYWXGDUS2OF5/VCNP4125_FAV.JPG PHOTOS 31 aims a bite at 29’s flank as the other dog retreats. This would be scored as a territorial dispute and chase (TD, C) but with no fight (F), as the two prairie dogs did not roll into full-contact fighting. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493932403119-VPAJ9LEW5XXFJHTO2FY1/VCNP3886.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493932440777-VGFBX6CYX1PYJWBJWYVN/VCNP3912_FAV.JPG PHOTOS Both males jump away from each other upon breaking a hostile kiss. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493932495198-2LKOF0BX7DAJCN99N1RY/VCNP3916.JPG PHOTOS 29 and 45 go airborne in an intense contact fight. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493932511343-MHJEJJ43309D5BEYFJWK/VCNP3917.JPG PHOTOS The two dogs break apart. Contact fights usually only last a couple to a few seconds. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933111548-VHU0DTACH3XH3LJ9T9JP/VCNP6552.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933212683-NV6C2QMFDEY4W4UHGYQV/VCNP4878.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933227663-9E5JIX0UCNITW7NSTFYA/VCNP4886.JPG PHOTOS A full contact fight. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933267892-G8OPNC13I1DQCFQYDTHG/VCNP4887_FAV.JPG PHOTOS The fight breaks apart. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933336830-3HXF9OHB1BQDQLW82AG3/VCNP5193_FAV.JPG PHOTOS A full contact fight occurs. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933391894-SAAPLE0MYT8EXOM0W7WQ/VCNP5195.JPG PHOTOS A lucky still shot captures Triple Stripe biting 51 in the face as they fight. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933486094-N19WUSM5EBFJG3QUS0T5/VCNP5661_FAV.JPG PHOTOS 49 tackles R21. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933511943-9FAW2IOV2OHXNAY2WY0H/VCNP5663.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493933639449-6WJ8P1FQ0O8SJTVLTROL/VCNP5522_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493934060596-EC3E3C08NCHPCQOTQI2L/VCNP5554.JPG PHOTOS The two females again break into a contact fight. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493934156418-70L9AIT0J5NCPDJIXLVA/VCNP4426.JPG PHOTOS On this day, WA9 had ventured too close to the burrow Wetsuit had claimed. Though these two females live in the same clan and are blood related, after they have copulated they become defensively aggressive over potential nursery burrows. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504552617080-MEQ4CMRNFZYLEYA1S2Z1/VCNP3916_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504552627273-PJAFDL0L6JWL0N2HZD38/VCNP6552.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504553316710-ORP222FOFCIUXD3I72O1/VCNP9212_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713053925-27LFGBKJMZ9GQT8XX01U/IMG_2388_FAV.JPG PHOTOS ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713055438-NTIW60HM8NOJ4KLAH4EN/IMG_3190_FAV_TD.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713068893-2NAD95ILR8VK3772Y74W/IMG_3242_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713068558-3C4FQDUN2TUDY5VJKKPU/IMG_3245_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713073781-BHAU8MC07ZFS3BMD83QQ/IMG_3317_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713078885-SKIPMJDWJ5M07CJ449TY/IMG_3338_FAV.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713088460-V1232WMF7YGSCLXVBEHJ/IMG_3410_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713085154-WE210T99WRLSIGHNYND6/IMG_3415_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713101756-W63D0L0889X8T0MHL69L/IMG_3453_FAV.JPG PHOTOS ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713096968-5LCK7LISYPDHUMA6NQNL/IMG_4218_FAV.JPG PHOTOS ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505765887641-1F9J3805NKXGQUMK2WE9/1219621.jpg PHOTOS - battle gallery https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505765622691-D65JL50IO0TQPUSI98L1/1219621.jpg PHOTOS - portraits in the prairie https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507160724836-2V7HCQ7LOPCRGKJENI5D/VCNP1096_NEW.PNG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507310959995-1HYHAB1GUH9KPIETZ02D/VCNP2284_1_NEW.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504386473801-9G6VUA50GLGXRESKF9H9/VCNP5724_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504383825785-P6RMWA80GZZ2JPVBXLN8/VCNP3128_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505835243082-GOK6C93MA5ISKPY3TEDK/EMBOND_5509.jpg PHOTOS Photo by Elaine Miller Bond (www.elainemillerbond.com) https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504387282049-I48KV3B6Z4IITU2UQBCM/VCNP4241_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504383830752-PZED1GGCA7LV5SIP36IB/VCNP3383_1_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504386401705-GF28MIYVZTSB283ORMGU/VCNP2347_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504386390590-WRLU8WY82PFFM6XLSEZG/VCNP2446_FAV_alert.JPG PHOTOS ©MRR 2017 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https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504549790751-5QNW6D7MBYH3DRYHCS7X/VCNP8140_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504387256073-TRCTTPUQY7B5TQYE9VKV/VCNP7939_1_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505921737521-KJEA7BSDX86WEJIXART1/VCNP6414_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504550893965-XZA5I9EMS6SAH3WWDIQN/VCNP8015_FAV.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505668836546-96IQQMIB7FJA8W0DYTDS/VCNP8414.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505668846448-AEK4RZHJH8M8NTMGUZ39/VCNP8417.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507310962771-Z0ZJPQCIDX405MPQ3JLM/VCNP1742_NEW.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507910902114-SXS9X780QD5E7PDJNL2R/VCNP4403_FAV_TC.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507130185416-W40ZI7OAUFDLVSVNV289/1219621.jpg PHOTOS - Eyeview GoPro Captures https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129155763-6KYS3EGEFN2NBUZZYB9S/GOPR0804.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129092539-JATA7DIFGEQM5C4AY4XK/G0010295.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129107628-70V6G8VSF1HR2D5UMP7Q/G0020823.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129107764-YYG0G0RIGG5FB9SW4IOT/GOPR0105.JPG PHOTOS ©MRR 2017   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129150677-78R9W8QM5KN4C4GOAC7A/GOPR0126_alert.JPG PHOTOS ©MRR 2017   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129120469-N76CPMPTVN7X8GDAF3VS/GOPR0179.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129129148-8DCUA30E8O7KWW2PHW4O/GOPR0500.JPG PHOTOS Snow-covered mornings. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129144133-5HM9571M257QQ52X71MV/GOPR0799.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129166661-Q8P3MHVAWTORZ53Q9LCC/GOPR0851.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129173922-F1H4OTGYXBW8O7N8GAW5/GOPR0854.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129212204-5W6CJF2QL6KOR3E2N3V3/GOPR0548.JPG PHOTOS Big space, little prairie dog. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129652371-7PH93N9FF810YWO72PLV/GOPR0679.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129635317-IRWDEXA61TGNKGHU7CDN/GOPR0685.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129649042-ZE03L3FS7I7BQQSLU5E1/GOPR0693.JPG PHOTOS Emerging for the first time. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129665338-8HWNEJ05FPFP2V9KED18/GOPR0728.JPG PHOTOS Approaching dusk. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129672750-6DT68EEFVQSOZXFJCTMY/GOPR0758.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129685974-5RFCR25DC77ZTWWS0AQJ/GOPR0784.JPG PHOTOS ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507129684967-50CPKDO9SC8H6O9K6FCD/GOPR0864.JPG PHOTOS Surrounded. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504711594600-YMZCG7BUJFZJ3PTF3JNQ/IMG_3317_FAV.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504711656065-S1TXO82F6VB3HJ3W7JO5/IMG_2013_FAV.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504712671710-K6IOIJ4EDEL1P0VDAENQ/VCNP1736_FAV.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731410251-LQ4WMINV00ECJ2SFMA6D/IMG_4285_FAV.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839143944-FUFHVN5E9A0H4O45R1Q4/VCNP6136_FAV.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507130683815-KUKI1JXAE45RZ3KG0KOF/GOPR0804.JPG PHOTOS https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507160946230-2TR0Z52WL2SIDT0CKNTK/VCNP1096_NEW.PNG PHOTOS https://www.prairiedoghoogland.com/the-project-index daily 0.75 2018-02-28 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495580911468-XZZCPFOFAGGOJGWDIZ0K/IMG_4555.JPG THE PROJECT Squaddie Rashidah marks her flagging tape with the burrow ID where this prairie dog was captured. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494616035004-6THSEO2VJUHN6Q4JWX5K/IMG_3387.JPG THE PROJECT Trapped prairie dogs wait in the marking van for handling. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494616028066-YLHKI1H58F8IF8JOQUGL/IMG_2390.JPG THE PROJECT John checks a marked prairie dog for eartags. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504888190821-X48VJRZ62VDJXW6X3XHM/VCNP5734_1.JPG THE PROJECT Weighing an adult in the handling bag. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504888218573-3RJ21M7S8SHOBHU1QDOJ/VCNP5946_1_FAV.JPG THE PROJECT Weighing a juvenile is easier in a plastic bag. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504888204722-CJOOUM7LKX75B5FUE98P/VCNP5784.JPG THE PROJECT The handling bag allows for easy inspection of reproductive condition. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504888219425-XP0LCVT909KQB55BU7V3/VCNP5745_1.JPG THE PROJECT The earpunch takes a small amount of tissue. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504887854048-ZEW2S31F0STC2LHX8X9P/VCNP5715_1.JPG THE PROJECT A closeup of an eartag. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504887862087-2PVWDW45NKXZ68T3M1BA/VCNP5726_1.JPG THE PROJECT Tissue samples go into individual vials for each prairie dog. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504888238406-DRXBM1G54Z9P2U1GM26J/VCNP5967_FAV.JPG THE PROJECT Eartag and vial numbers must match. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507128854433-6GC8SIJCM61YM1T76HAY/GOPR0122.JPG THE PROJECT A handling datasheet. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494616033485-5T21XM4LA0URESZOVYXE/IMG_7741.JPG THE PROJECT Marking Male 41 with Nyanzol dye. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504888232398-WGH63CY93ITC9LW0KDVG/VCNP6068.JPG THE PROJECT Juveniles hold still with only a small amount of pressure around the head. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505777681585-PW60XIL6QHJAGH50R97O/VCNP0763_FAV.JPG THE PROJECT An ultrasound wand and gel. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494615277921-5C4BNDXSBZL5XN2R2QLI/GOPR0425.JPG THE PROJECT A prairie dog's view inside a surrounding. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494615288754-YPJHTJD3Z6SMMZGZUNAR/IMG_2388.JPG THE PROJECT A prairie dog waits for handling. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495580186476-5EN3P4EPBGFLSRTPWK83/VCNP3094.JPG THE PROJECT An escape artist climbs over a surrounding. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495580206513-Z0R9RCCUXEBIORU4XD7W/IMG_5154.JPG THE PROJECT A double-door adult trap and a single-door juvenile trap - both Tomahawk live traps. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495580227070-RXJWDIFVSBRN33YVOMR4/IMG_3280.JPG THE PROJECT Traffic cones are used to temporarily plug escape routes for hard-to-catch prairie dogs. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495580555553-T8T17XIF2ZRMFSURX5OT/VCNP2992.JPG THE PROJECT Squaddie Patrick and Denver Zoo volunteer Tim open traps at a surrounding. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495580238616-ZKVXEN4FX0SMGHKN8JBF/VCNP2275.JPG THE PROJECT John clears snow from a spread trap set and adds bait. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505671623865-YRB40OK3P17SRBQ29130/VCNP6954.JPG THE PROJECT On especially hot days, we cover traps with white cloth to provide caught prairie dogs protection from the sun. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495580564187-QDAN2RW6YLKY54P5PF7L/VCNP4326.JPG THE PROJECT An unmarked prairie dog caught in a surrounding. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504288100317-N09GWEHKNUFHK0GW4LKE/IMG_7728.JPG THE PROJECT Squaddie Marlin setting spread traps. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504552358807-BPWD67FBRYSM9BCAOS0B/VCNP2661.JPG THE PROJECT A large male caught in a trap. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504552368023-RSNZ5FSI4RHR2QAR79QI/VCNP4839.JPG THE PROJECT Sometimes the prairie dogs refuse to get caught. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505777376806-RY57BFHII2G14103SYEH/11.45.15+hrs+__%5B0001277%5D.jpg THE PROJECT The spine of one fetus and the skull of another. ©John Hoogland 2015 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505768215127-OW6J9GQ1IRSIT98VI0M3/VCNP5998_1_FAV.JPG THE PROJECT - Purpose and methods https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505777410731-TUSD6AIZ772KH536EGVT/15.28.25+hrs+__%5B0001284%5D.jpg THE PROJECT A clear spine. ©John Hoogland 2015 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494614355534-PX95TBD1I549HD9C52Q6/IMG_3379.JPG THE PROJECT The inside of an observation tower. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494614340393-N1YFAY8LJ2OFK3Y50UCU/IMG_3090.JPG THE PROJECT The observation tower throws a long shadow in the early evening. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494614335405-73EDH5UVI4MEKWMBZ4NR/GOPR0202.JPG THE PROJECT A closeup shot of a prairie dog at his burrow shows an observation blind in the near distance. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494614367272-OAF6C2X8SCVTB6LX6DTP/VCNP2994.JPG THE PROJECT Observation blinds are positioned at key locations across the colony. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494867729958-SOVAQEB5ZNS4AAKIOPVP/IMG_7761.JPG THE PROJECT Squaddie Katie demonstrates the cozy fit in our new observation blinds. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504288255599-HJ6ANFGJ8ZXLQHZSLBPX/IMG_7805.JPG THE PROJECT Sometimes we work out of our cars to get a better angle on activities, whether for photography, data collection, or both. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505870822762-0QSBCW48H1SARMVIQJ8I/GOPR0101.JPG THE PROJECT As longs as the dogs are out, we are out, inclement weather notwithstanding. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1517586912041-2SNNAEX2VA08ACT5BMBE/IMG_0672.JPG THE PROJECT On quiet days, we sometimes turn to art. ©MRR 2018 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505768174072-1KPRQGRC6GPFV1XPA4BZ/image-asset.jpeg THE PROJECT - sTUDY hISTORY https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505771730126-N5QCDBQDD8VTV5XH2BLS/IMG_5003.JPG THE PROJECT - mARIANA rIVERA rODRIGUEZ collaborator https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505689032114-URGUNQAK5UMVJO3SUY0B/IMG_7665.JPG THE PROJECT John and squaddies. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505770126207-4PFTP7OCLG6981CAIMNI/IMG_7740.JPG THE PROJECT Helping John in the marking van. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505770143580-1AG914K65MIMSNX1CT1U/IMG_7748.JPG THE PROJECT Marlin. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505773088210-F7IAPTZO1PCVYYYVSYZQ/IMG_1663.JPG THE PROJECT Adjustments in the tower. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505689028171-QMYS5PL31WYIVV4V29C4/IMG_7676.JPG THE PROJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505770132741-NBS8ADRGXKV7053D49LZ/IMG_7855.JPG THE PROJECT Patrick. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505770137917-X73GG7SD76QDZXL8TETI/IMG_8451.JPG THE PROJECT Katie. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505773354251-0D7H0CFXSD43OYM6U7M2/IMG_2747.JPG THE PROJECT A squaddie in one of our older observation towers. ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505770159480-NNBLE20AIQF0RPI6G6JH/VCNP4471_1.JPG THE PROJECT Carly, Dennis, Katie, John, and Jimmy. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505770233413-EBL0DX85PP2MB2213EDO/VCNP2284_FAV.JPG THE PROJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505772670791-DB5TXFHPP8W04SHARPQF/image1.JPG THE PROJECT Marlin and a snowprairiedog. ©Marlin Dart 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505772837788-9SLC27HRF3LKIENA7RIV/VCNP7697.JPG THE PROJECT Patrick setting a surrounding. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505772844825-N3ZY0LCR9RW8KKN661GF/VCNP1683.JPG THE PROJECT The squad arriving for the morning. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505774495529-XTQF4EBBJ89CQRT78IWM/VCNP6154.JPG THE PROJECT Dennis with a juvenile. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505773035234-0ZHQ46FL0XME021WJXD2/IMG_1556.JPG THE PROJECT Observer in an observation tower. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505772899586-AXCUD1LS852ZZ5FSY1PC/VCNP2995.JPG THE PROJECT Patrick and Denver Zoo volunteer Tim. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505773785930-9IDZV82B8LVJHZ286153/IMG_7756.JPG THE PROJECT Us and our binocs. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505773706971-5UM4ZFSH8FTULAQBMC0A/IMG_7691.JPG THE PROJECT Walking away at the end of the day. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505773785222-LDJI1VPDJXP2EG52X04V/IMG_7761.JPG THE PROJECT Squaddie Katie in a new observation blind. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505774401858-YL8Y4VMX0JEX9KCURBEM/VCNP6269_1_FAV.JPG THE PROJECT Carly with a juvenile. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505771548894-FJ3EOGE82GWIVG5GC350/VCNP4471_1.JPG THE PROJECT - The Prairie Dog Squad field assistants https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505768317287-OCQNLNFLFAT8CD49OD2J/image-asset.jpeg THE PROJECT - THE RESEARCH TEAM https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505771317732-ERUMN4ZZPE8C50XS6RLQ/VCNP2280.JPG THE PROJECT - John hoogland principal investigator https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496286878351-UDB8EDVIOJA2PGQ38WX4/image-asset.jpeg THE PROJECT Male 42 licks his penis after mating, a telltale sign that a copulation has occurred. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496265732523-AY8CQVDH7YTVHICB3U8B/IMG_4578.JPG THE PROJECT A prominent and visible burrow mound with a large entrance. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496265747663-NM2B834C3266V566KEV6/IMG_4519.JPG THE PROJECT A prominent mound with a smaller entrance. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496265742054-TUKTE0G6FV5E6G5KQUAI/IMG_5605.JPG THE PROJECT A tiny burrow entrance with no prominent mound. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496287052986-3GX8GPOKF8K6R6E9OP8F/image-asset.jpeg THE PROJECT Not even the safety of a burrow can save juvenile prairie dogs from a persistent weasel. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507835135435-E7PLDJSE0T6XDB1SIJPO/VCNP7407_FAV.JPG THE PROJECT A long body and stout legs for underground tunneling. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496286531831-4WWPFRAMBH4LO2SJND6T/image-asset.jpeg THE PROJECT A mother transfers her recently-born baby from one burrow to another. This juvenile is still pink and hairless, its eyes still closed. Its birth occurred underground and it will be a few weeks yet before the litter is ready to emerge on their own. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505768276029-SAKOM6BVBR7S2Q0NRA03/image-asset.jpeg THE PROJECT - aBOVEGROUND cUES https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496286296733-3ANSII9BK4E9I7ZIXRBM/image-asset.jpeg THE PROJECT The nursing rings around this mother's teats are visible when she stands on alert. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504550397414-T0SDEH9U58D900C10ZR7/image-asset.jpeg THE PROJECT Parting after a hostile kiss. ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504308147244-Y7YRG4QEI830W0SZQ90K/IMG_2388_FAV.JPG THE PROJECT A territorial dispute escalates into a contact fight. ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504248035113-0J4Z687EZFEDZPIZTGGR/VCNP9249_FAV.JPG THE PROJECT RACR works her mound, removing rocks and smoothing dirt. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504241936686-O9GMR96KTKI4RVJRW63X/VCNP8439_FAV.JPG THE PROJECT Male 36 leans over to lick his penis after copulating. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504244879412-Q5ZUZTS3ZUQ9ZXO2UJOU/IMG_8313.JPG THE PROJECT A predator-pull alarm-call experiment datasheet. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504308978117-DEKPUBSHH9EM34FTUG0D/VCNP8051_FAV.JPG THE PROJECT Male 2 comes up behind female Wide Apart to sniff between her legs. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504232923910-2FBV9KQAHLR7HQNH1HDK/VCNP5661_FAV.JPG THE PROJECT Two males in a contact fight. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504219351336-G7XL9T6OH0I44BJY182Z/IMG_7782.JPG THE PROJECT Arrows from one line to another indicate the two interactions are connected. On this sheet, at 3:30 4 chased 18 into a hole, 45 witnessed this abuse and, not to be outdone, then chased 45 farther into his hole, at which point 4 ran away from 45. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504219323502-BW54471HABYIXRKW6GWT/IMG_7792.JPG THE PROJECT This page shows several "BDs" which is code for Both Down. When a male and female prairie dog are underground together for longer than 15 seconds during the mating season, we call it a BD. BDs that last longer than about 5 minutes are usually critical copulations. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504219513445-1QT4QVLHHWAXN0BDGGV9/IMG_7852.JPG THE PROJECT This sheet shows typical male behavior during the mating season. Here 24 spends a considerable amount of time pursuing females to check for estrus, and chasing males out of his area. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504246004350-5KRF1QPQ3S5C505XGTY3/IMG_8257.JPG THE PROJECT Sometimes it's impossible to code important information onto a single line, and we may find ourselves having to write a "story" of a key observation or occurrence. In these instances (including infanticidal events, predations, major injuries or deaths), the more detail the better. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504228363957-C2JUBC3L8JEXDOFIS4AH/VCNP2487_FAV.JPG THE PROJECT A friendly kiss; an adult prairie dog kisses a baby to say hello and determine if the offspring is hers. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504307886716-T0EPT9Q1R3NKINBII671/IMG_1725.JPG THE PROJECT A mother brings nesting material to her nursery burrow. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504245724062-DO1RIF2FCIT54W2FF8ZB/IMG_5171.JPG THE PROJECT A Wakeup/Bed and Baby Census datasheet. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505749761154-9GVHF5NWANNKMWTQ6M6S/IMG_7733.JPG THE PROJECT - cODING bEHAVIOR https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504229536691-9U6HBFEON2CBD0M3BAKK/VCNP5477_FAV.JPG THE PROJECT Two feuding males nearly make contact during a high-speed chase. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504308318085-S01B51WCCEZH3HFO2L4Y/IMG_2559_FAV_C.JPG THE PROJECT A chase. ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504282813220-JNKNSRI989TDSGOL6JNG/VCNP8844_FAV.JPG THE PROJECT Using her front paws to adjust stray strands of grass in her mouth. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504226885862-TOKRH3C8WJ6L04NTGP47/VCNP7544_FAV.JPG THE PROJECT A hostile kiss between two males during a territorial dispute. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1492449431964-KQJL0W202EEG3AUPBOLC/VCNP4887_FAV.JPG THE PROJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493166097397-55M8SZTDY2V4QUWYQ3ZJ/VCNP1557.JPG THE PROJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1494545794472-NSHTRFZQF4U977B73RIX/VCNP3007.JPG THE PROJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495746135416-82EXHTFT3J7L77RK5BYG/GOPR0126.JPG THE PROJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504205935394-PVX0O073E4YEF4VCGSF2/IMG_7733.JPG THE PROJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504384969846-I8WTY8U7EPEFG5T8HVVI/VCNP5998_1_FAV.JPG THE PROJECT https://www.prairiedoghoogland.com/our-subject-index daily 0.75 2017-10-12 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495581892528-VXBQMEF0EX9FJK7WRJ3C/image-asset.jpeg OUR SUBJECT The geographic distribution of all five prairie dog species. Image from Conservation of the Black-Tailed Prairie Dog, edited by John Hoogland. https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504913727396-WFYS1BJ7ES6WFHLBBEQ4/IMG_3394_FAV.JPG OUR SUBJECT Snow is common across the prairie dog range. They may do their best to avoid the cold and wet, but once the mating season has begun, not only a spring snowfall can keep the prairie dogs underground. ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839642844-OI0S9ORQL6YVJSY1DZKE/image-asset.jpeg OUR SUBJECT - NATURAL HISTORY https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504190431663-2HPI5ILO3TFPKIIZFF5V/VCNP4520.JPG OUR SUBJECT A Gunnison's prairie dog (Cynomys gunnisoni) in New Mexico ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504190299665-T6NZGCH8VIJ2QSG7J3OW/IMG_4214.JPG OUR SUBJECT A white-tailed prairie dog (Cynomys leucurus) in Colorado ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839916676-POI6XEYPD7QOABSQLEHO/VCNP5578.JPG OUR SUBJECT - CONSERVATION https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839692796-ZHFS52B75X2M9U088T2O/VCNP6554_1_FAV.JPG OUR SUBJECT - COLoniality https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942709262-JGEJS27APD369WQJPG4Q/VCNP5659.JPG OUR SUBJECT A territorial dispute (TD) between males R21 and 49 begins with a hostile kiss. TDs typically begin with mouth-to-mouth touching and sometimes flank-to-flank posturing. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942696242-87UHELGFDLXVTEPV0WHX/VCNP5660.JPG OUR SUBJECT Breaking the hostile kiss, R21 leaps out of the way of 49's snapping jaws. When a contact fight occurs, prairie dogs use their teeth and claws to cause injury. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942730494-OJ0NEJXP832LP6U67RGM/VCNP5661_FAV.JPG OUR SUBJECT 49 catches R21 in a tackle. Prairie dog fights often look like a ball of fur mid-air or on the ground, and are over in seconds. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942724263-BN9QVUGAVPQ6GAN10XQG/VCNP5663.JPG OUR SUBJECT During a contact fight, the two prairie dogs will roll over each other trying to gain the upper hand and deliver bites and blows. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942769939-KQUOT4EOBFSVB15TXGSS/VCNP5664.JPG OUR SUBJECT 49 and R21 continue to roll, kick, and bite. These pictures were taken in a burst, fractions of a second apart, demonstrating the speed at which these animals move. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942743460-SWIOD6VX7OEC8JWP1Y5J/VCNP5668.JPG OUR SUBJECT The two males break apart. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942756147-6BYZ6NNE3E3RVP0APNSA/VCNP5669.JPG OUR SUBJECT R21 immediately twists to his feet so he can flee. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942768973-ONK9N204ZUZOYE3JVUKJ/VCNP5670.JPG OUR SUBJECT R21 pushes off to run away. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493942780807-TMDRE4VQCS5PFTKS60B8/VCNP5676.JPG OUR SUBJECT 49 is in hot pursuit after his tussle with R21. Most fights end with one of the prairie dogs running away and the other either giving chase or running in the opposite direction. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504191456603-TFZCOCRVURNEDE5PGTV3/VCNP3001.JPG OUR SUBJECT All the members of this subcolony/ward are on alert. Juveniles can be seen in the foreground at burrow N5. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493941706916-AKNYEQ8XSEAE8YGPLV8J/VCNP9387.JPG OUR SUBJECT R39, a yearling male, inspects RR0's teats, though she is not lactating. It's possible RR0 is R39's mother. If so, she would have given birth to him the year before. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493941743477-GC0LNPFKXTQRF7J2ZY6O/VCNP9388.JPG OUR SUBJECT After inspecting RR0, R39 somersaults playfully. Whatever RR0's relationship to him, the adult female is certainly kin to R39 and the yearling male will disperse before he becomes sexually mature. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493941741023-2F24XZGH3T0FNN9D0KGQ/VCNP9520.JPG OUR SUBJECT R45, Head 2, and 90 are all kin. Females Head 2 and 90 will likely spend their entire lives in the same ward, where yearling male R45 will disperse. Adult male 36 would have come from another ward before dispersing and establishing himself as a dominant male in this area. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731059001-3YDC050VPKC3ENZS7O52/IMG_4285_FAV.JPG OUR SUBJECT ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917554578-VOW15EXTO2JP3BVXV0S6/VCNP1862.JPG OUR SUBJECT Males 29 and 45 have a territorial dispute. Many females have emerged in their area, and they will dispute through the mating season, often fighting. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917533162-9IVA4TUCNTPP5WCQRWZ6/VCNP2678.JPG OUR SUBJECT During the mating season, adult males are evidently sexually mature, displaying pigmented testes and a distended scrotum. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917553818-7T1OAZ9PCISS58XFZEMU/VCNP3152_FAV.JPG OUR SUBJECT 45 stands over Wetsuit at her burrow, anticipating her estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493917730520-1NG1V0JROJ65ATQRGPPJ/VCNP3191.JPG OUR SUBJECT 45’s pursuit of Wetsuit is relentless on the day she goes into estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918059322-79HWV5RC9Q9VSFYFS9IP/VCNP3750.JPG OUR SUBJECT 45 brings nesting material to a burrow prior to mating with Wetsuit. Though not common, this behavior has been observed in mating males. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918020907-WNGXLBCZWOAWDZNGW1W3/VCNP3777.JPG OUR SUBJECT 45 lets out a mating call at the mouth of the burrow where he has chased Wetsuit. The mating call is sounded before and/or after mating, and if not for context can easily be mistaken for an alarm call. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918626633-Q1NJMLXCGFOPH8E25WQV/VCNP4553.JPG OUR SUBJECT 45 chases and herds 50 into her burrow. We call this a Chase Herd (scored as CH on a datasheet) and it is one of the most common interactions we track, occurring between males, between females, and males to females, though rarely females to males unless that male is a yearling. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918785742-CWHW5COZHTNXM4LLNKYC/VCNP4578.JPG OUR SUBJECT Generally a couple to a few days after mating, females will begin to exhibit maternal behavior, including carrying nesting material to potential nursery burrows. Gestation will last on average 28 days. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493918748561-CZ55GUCH8LOPTN5XM7D7/VCNP4600.JPG OUR SUBJECT After mating, 83 licks her vulva. This is one of several aboveground cues that signify a copulation has occurred. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919327047-PP8F7MSUKSD094FV4PV2/VCNP5960.JPG OUR SUBJECT Despite this being her day of estrus, 83 is not ready and rebuffs 45 for now. On their day of estrus, females are only receptive for a few hours. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919486184-S96CE7WO4XNHWVN5B94W/VCNP6300.JPG OUR SUBJECT 24 has a very late wakeup and licks his penis (scored as LP on a datasheet) immediately upon emerging, indicating he spent the morning copulating with a female. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919480313-ZCEMWQW1UOU9WVX0H4MV/VCNP6379.JPG OUR SUBJECT Having already mated with her, 24 tries to keep 6 Stripe underground, chasing and herding her into her burrow to keep her away from other males in the area. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919665906-YX3T8QO020XS5FB9EE7R/VCNP6429.JPG OUR SUBJECT After a male has had his Critical Underground Consortship (CUC) with a female (in which insemination occurs and which is characterized by disappearing underground together for many minutes up to hours), he usually does not go down with her for another copulation. Here 24 and 6 Stripe share a hostile kiss as he tries to keep her at her burrow, having already had their CUC. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919686605-5RH6S55J8U44QUB8S3A6/VCNP6446.JPG OUR SUBJECT 24 will relentlessly chase 6 Stripe to keep her away from other males. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919806245-CYS0VC1CN5RNEAH7DCKS/VCNP8153.JPG OUR SUBJECT R21 chases 49 out of his territory on a day when 5SRS, a female in the area, goes into estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493919824286-4U2DSQ43NW9FW4X9IBZ7/VCNP8257.JPG OUR SUBJECT Some time after his first check, 2 comes back for another sniff at Wide Apart. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493920128923-4RKOYK8D09XTIPSCFQET/VCNP9594.JPG OUR SUBJECT Male 36 investigates female 90. Gunnison’s prairie dogs exhibit sexual dimorphism in body mass, and here you can see how much larger 36 is than 90. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493920188437-VS8HVCU3K4QAGQ51F4PZ/VCNP0027_FAV.JPG OUR SUBJECT 36 continues to pursue 90 to check for estrus, and she continues to elude him. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493920201208-ON7IG5WRXV19GBLBDLCD/VCNP0112.JPG OUR SUBJECT RR0, a female who has already mated, brings nesting material to her burrow as she begins to show maternal nesting behaviors. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551737683-82W469V6LK1DDIBRNVSX/VCNP4760.JPG OUR SUBJECT A female runs away from a male trying to sniff her. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551777256-Q6QSL86O0495XI5DC35H/VCNP4615_FAV_LV.JPG OUR SUBJECT Female 4 Stripe licks her vulva on her day of estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551785372-BKR2REUDRHPB8C2KK4XL/VCNP6403.JPG OUR SUBJECT Female 55 runs from male 36 as he pursues her. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551839458-N5RXHIPT173IA2DUISG4/VCNP7882_MC.JPG OUR SUBJECT A mating call from male R21. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551815115-9F0MNLXSFBKQRRO59BY0/VCNP7921_MC.JPG OUR SUBJECT Another mating call from R21. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551833827-LDFXVBXWGX7OFVWE4V4I/VCNP8256.JPG OUR SUBJECT Male 2 comes up behind female Wide Apart to check her for estrus. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504551870261-W8Y13KUHQYFY2UWZYVZ1/VCNP8439_FAV_LP.JPG OUR SUBJECT Licking after copulation. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713772960-51SZ8MLRYRTUQ0ZF1OID/IMG_2384_FAV_MC.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713790127-T7KQL76O4KA02L65EGCS/IMG_2375_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713780642-J4N5UNCCDPTPUITGQHPZ/IMG_2438_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731072258-76O9DQYTSZ7UK9IESKTI/IMG_4299_FAV.JPG OUR SUBJECT ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713800949-OV6WF3FDDLRXBK2M3E3P/IMG_2439_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504713797881-12PYTA9NVUCT97Q09XAH/IMG_2440_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730766222-KKSAJE2L3WF9SQI5JZ3T/IMG_2603_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730765541-FISPU7RKCP9LCXIPP42T/IMG_2640_FAV_MC.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730793394-7L9173U26PKHJKKZ4QR4/IMG_2644_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730777117-VPPWORPVSA1E7OC1Q0XS/IMG_2647_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730789551-UTIKC0EH9KJVG2TQP2RV/IMG_2696_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730936930-JBNEYMWJRRZNJCRYTOBO/IMG_2707_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730931179-QRZN9MYUMPBVAE00PCL0/IMG_2735_FAV.JPG OUR SUBJECT ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730951486-4RFIZSINOQJH9LLIXLCB/IMG_3067_FAV.JPG OUR SUBJECT ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730946815-7EK9U56BEYS7K1AE5WCO/IMG_3324_FAV.JPG OUR SUBJECT ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504730958266-82OCTOEEP5OQFGISJ2JC/IMG_4276_FAV.JPG OUR SUBJECT ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731068539-IJ0BAH29SHA9LTAZUBMD/IMG_4303_1_FAV_MC.JPG OUR SUBJECT ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839735410-EP6BWMQRJE77EDQPA14Q/IMG_4299_FAV.JPG OUR SUBJECT - THE MATING SYSTEM https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839775754-BWZWV8MYDWDMF9HTW846/IMG_4893.JPG OUR SUBJECT - INFANTICIDE https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496275803413-NLHGHKEFB19FNKS7EDNO/IMG_4795_FAV.JPG OUR SUBJECT Here male 29 cannibalizes the offspring of mother 79, after killing the juvenile in its burrow and carrying the body aboveground. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496275792053-G7XX7DZ30JFJA417FPRQ/IMG_4800_FAV.JPG OUR SUBJECT Male 29 continues to cannibalize the carcass of the juvenile he has killed. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496275799674-SBPETCKRFII9QSAAP5KW/IMG_4804_FAV.JPG OUR SUBJECT 29 bends over the body of the juvenile he's killed as he feeds on the carcass. This cannibilizing occured very near to the victimized burrow. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496275806978-APLMMTH1NYS8RVSQT8QJ/IMG_4830_FAV.JPG OUR SUBJECT Male 29 looks up to survey his surroundings, standing over a juvenile he has killed and partially cannibalized. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279687222-OMP4MGOMJ7BG4DMQRUW6/IMG_3911.JPG OUR SUBJECT The relatively longer black-tipped tail of the Wyoming ground squirrel can be used to differentiate it from the white-tailed prairie dog. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279694506-HVA6H85X9ZMQGZZ9FOM0/IMG_3916.JPG OUR SUBJECT Wyoming ground squirrel juveniles look nearly identical to white-tailed prairie dog juveniles. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279827555-P1HUWNIII8LWD0PXABST/IMG_3783.JPG OUR SUBJECT Female white-tailed praire dog RACR (Ring Around the Collar, Rear legs) faces off with a Wyoming ground squirrel. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279873832-355CN50Q1H41C1N1U09W/IMG_3789.JPG OUR SUBJECT A Wyoming ground squirrel mother defends her baby (background) from RACR. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279910894-2V59RYO6CPBDEBBV7KHS/IMG_3813.JPG OUR SUBJECT Female prairie dog RACR leans over the body of a juvenile Wyoming ground squirrel she has just killed, while the agitated mother looks on. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279658451-RA0X8NHV7MRJ0CZD9MCZ/IMG_3901.JPG OUR SUBJECT Prairie dog male 42 leans over a juvenile Wyoming ground squirrel he has just killed. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279667693-07K4I9KRF2JXHD29GY67/IMG_3904.JPG OUR SUBJECT Blood is visible on 42's face from killing a juvenile Wyoming ground squirrel. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279707732-7Q7FI9HG507RA06P9UH9/IMG_4479.JPG OUR SUBJECT Prairie dog 2RS (double Racing Stripe) investigates the carcass of a juvenile Wyoming ground squirrel she has killed. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279752096-F0HY4TRITREAUIU3CQTG/IMG_4465.JPG OUR SUBJECT The fresh wounds on the carcass of the juvenile Wyoming ground squirrel are evident. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279745222-NXMR9OT1BFFCABBJBGLS/IMG_4473.JPG OUR SUBJECT 2RS surveys her surroundings standing over the carcass of a juvenile Wyoming ground squirrel. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496280711115-2GAM5H36863G092EHWK2/IMG_4638.JPG OUR SUBJECT A lactating female prairie dog holds a dead juvenile Wyoming ground squirrel in her jaws. This image was featured in the National Geographic article featuring John and Charles Brown's research. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496280696176-0XHD1CHJMM0Y2A5VODX7/IMG_4643.JPG OUR SUBJECT The prairie dog makes sure the juvenile ground squirrel is dead. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496280707439-2U5IOYYZZ6TP96RKMU66/IMG_4651.JPG OUR SUBJECT The carcass of a juvenile Wyoming ground squirrel lays abandoned. White-tailed prairie dogs do not typically cannibalize their ground squirrel victims. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496279955320-5IFCFF9XJG4JTLZ5S3KC/IMG_3836.JPG OUR SUBJECT After an attack by a prairie dog, this Wyoming ground squirrel mother transfers her surviving offspring to a new, potentially safer, burrow. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1496280981983-V1YJIB207QGGBCVQR0BC/IMG_3931.JPG OUR SUBJECT A female Wyoming ground squirrel cannibilizes the carcass of a dead juvenile. This behavior was not well-understood. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504550173197-II21NWYHEP055OAOO32X/IMG_4660_1.JPG OUR SUBJECT Carcasses are examined and photographed for the record. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839809591-NAS6L7BP77VFW4QE78DM/image-asset.jpeg OUR SUBJECT - INTERSPECIFIC KILLING https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507155557442-85VLO0HD4KHGE4FH22TU/IMG_2384_FAV_MC.JPG OUR SUBJECT A mating call before copulation, with the estrous female still aboveground. ©John Hoogland 2009 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505842709568-1J5VQX4PB2WUSJ1W55K6/jumpyipfigure.png OUR SUBJECT Figure from W.J. Smith, S.L. Smith, J.G. Devilla, E.C. Oppenheimer. 1976. The Jump-yip display of the black-tailed prairie dog (Cynomys ludovicianus). Animal Behavior 24:609-621 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507155753678-3TMV1CCFIBMWBBF0DETG/IMG_4303_1_FAV_MC.JPG OUR SUBJECT A mating call after copulation. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504111818154-WE0HQBIOHGYF8OI7TWO9/VCNP3198_1_AC.JPG OUR SUBJECT Female RSBB demonstrates the tell-tale open-mouthed alarm call, pink tongue visible. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504306992889-I4MBQ3B7FL39RAHR7T3U/IMG_1569.JPG OUR SUBJECT A white-tailed prairie dog exhibiting the classic alarm call posture. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504111806826-DVIWWDJ42BAS1KFXXB2G/VCNP2639_AC.JPG OUR SUBJECT Though hard to see at a distance, this prairie dog is sounding an alarm call from the tall grass. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504111869565-Y0LOQ5W5A4EZ7SLXHB2P/VCNP2216_AC.JPG OUR SUBJECT Female Wetsuit sounds the alarm, likely upset by the traps surrounding her nursery burrow. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504307009015-8J9HVK47GL8XFO7E5T8B/IMG_1628.JPG OUR SUBJECT Like alarmed by a nearby badger. ©John Hoogland 2006   https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504111837858-RKSZ19WU5T61JJZCO1DQ/VCNP8111_AC.JPG OUR SUBJECT The lower incisors (front teeth) are visible during an alarm call. Though these calls can be given from a number of positions, up on their hind legs is the most common. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504111863396-TVBFLKRSRQV4DJMJKCS7/VCNP7454_AC.JPG OUR SUBJECT Many prairie dogs will run to the entrance of their burrow before sounding the alarm. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504111883130-NB43X9L376FUKDQWR0DT/VCNP5998_AC.JPG OUR SUBJECT Female Wetsuit stands tall on her hind legs to sound the alarm while keeping watch of the perceived threat. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504307004885-FE692Z5X24WU70BJMF4O/IMG_1744_FAV.JPG OUR SUBJECT This white-tailed prairie dog female shows off her wide mouth giving an alarm call. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507844368014-BQZ1KDE4WAMRLC5Y0MZS/VCNP8417.JPG OUR SUBJECT - Vocalizations https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504197572576-ZWKWQI0TD2NJ70G8CHA4/VCNP2453_TC.JPG OUR SUBJECT This is the typical posture of a territorial call. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504299834981-EW5JCF9TBNJK89IVFP2A/IMG_1377_FAV_TC.JPG OUR SUBJECT The white-tailed prairie dog will sometimes throw its head back in a territorial call. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504299847822-C5DY2MGO9L60Z9TD6SSU/IMG_1384_TC.JPG OUR SUBJECT And other times the white-tailed prairie dog will not throw its head back. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504197494195-EYLJA56WL0VFL6YJ5SLY/VCNP2953_FAV_TC.JPG OUR SUBJECT It's approaching evening and 45, the dominant male in his clan, lets out a territorial call. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504198419689-02K3M9PGMG6TXTQSS4S4/VCNP4403_FAV_TC.JPG OUR SUBJECT There are a few postures given for territorial calls, but a forward head and half-open mouth in an O is the tell-tale visual sign. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504197570585-YY8SM2ZZT0JG58072ZX7/VCNP3176_TC.JPG OUR SUBJECT 45 lets out a passionate territorial call mid-run. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504198466140-N9INMSWM0YGY5QW9URNZ/VCNP4647_TC.JPG OUR SUBJECT Though not as often as males, females also let out territorial calls throughout the day. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504299847254-6QQZTUD9HV0HPZK06BXP/IMG_1357.JPG OUR SUBJECT A white-tailed prairie dog territorial call. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504299857851-TXZ0D8Y6FM67T1SDRMZU/IMG_1695_FAV.JPG OUR SUBJECT The head-back posture can be extreme in the white-tailed species. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504198489859-FJ2WB2EWD79WM1TJJMWF/VCNP5032_TC.JPG OUR SUBJECT R21 stops mid-fight to let out a territorial call. A clear scar on his left cheek was acquired from one such fight with another male. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504198523956-P1XIBJMZSLG0KLFBVDVR/VCNP5084_TC.JPG OUR SUBJECT 0 (zero), the dominant male in his clan, spends all day during the mating season running around and establishing his territory. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504198541456-5M74I7K8X8LRCC3J6DKY/VCNP5097_FAV_TC.JPG OUR SUBJECT The classic O mouth shape can be seen here as 0 lets out a territorial call mid-run. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504198555619-8A2V2KB3H1MVVDRIRD0C/VCNP5155_TC.JPG OUR SUBJECT A classic territorial call posture. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504198566805-XZ6XHCDZOLPA97LGZ4V4/VCNP5757_FAV_TC.JPG OUR SUBJECT Snow-covered mornings see prairie dogs waking up late and slow, but the dominant male always comes out early and lets out a territorial call. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504199340197-HBT83D1CZHXREB4IYPF8/VCNP4472_TC.JPG OUR SUBJECT During the mating season, males will visit the burrows in their territory looking for females and establishing their dominance. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504299869300-JXJT055BQ9ZWRQRCUBNS/IMG_3466_FAV_TC.JPG OUR SUBJECT Another white-tailed prairie dog giving a territorial call without the head thrown back. ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504199383993-R1NNFDA7D43X794WT1PW/VCNP8130_TC.JPG OUR SUBJECT R21 did a lot of traveling as he attempted to secure his own territory, letting out frequent territorial calls. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504299878615-75E0VH646GESK61XZ53E/IMG_3464.JPG OUR SUBJECT Another head-back territorial call by a white-tailed prairie dog. ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504299884791-R055U83H7IKULVSR65SB/VCNP2062.JPG OUR SUBJECT This rare image shows a Gunnison's prairie dog throwing his head back during a territorial call. The head throw is not known within this species, but this particular individual (adult male 5) did it often. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505679565014-LU4WM7L1VRM0HXYFTZQH/VCNP4629_1_FAV.JPG OUR SUBJECT Mother and daughter. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505679321580-9B12I5BAB9WX2VGQEADN/VCNP3530_FAV.JPG OUR SUBJECT A transfer by carrying. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505679377834-QGFEA3N21V3S9JW0T99F/VCNP3356_FAV.JPG OUR SUBJECT A transfer by leading. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841026218-WP691GS7N9MN0FTUGE8Y/VCNP2738_1.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841040890-N30363E4BGSJGDZR5W5P/VCNP2739_1.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841078687-JHD5Q1EJ9KVQSQ3EYU14/VCNP2740.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841059017-TJEI3VTXD4F7ED5NQPYX/VCNP2741_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841077958-Q3Z1P3LNWKAYJYDDVGR8/VCNP3503_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841135808-4DRRGUKMTAS92Q8X5OUV/VCNP3504.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841099266-KQJVA55MIYEY94EW5SNO/VCNP3505.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841118840-DV4B4B1PUNX6BLS759NS/VCNP3506.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841137216-MUDXIKYP69N4H2FLCSFB/VCNP3509.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841199775-VWGVHQU61Y49AK3LEOUT/VCNP3512_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841156434-3BIW1I30T6MQZ2FDZZ58/VCNP3513.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841175780-BPA4PQ0S9KFZVXHQI68O/VCNP3516.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841194304-GO1DXEP28W0R58L0BGAM/VCNP3520.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841216819-9RU57P3KNDIXB665BLLU/VCNP4137_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841299881-RJPW3PRIJSYDZYV6QOOH/VCNP4138.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841259306-EIGMDHRSPEC9UBP06AH9/VCNP4141.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841275455-NHDWV0OI0L0JPQOUMQHG/VCNP4187.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841291300-Y7JQJOQWZNNT38YJAQ9O/VCNP4188_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841344336-P9TFPPKQVZSN57XKS47Y/VCNP4189.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841315481-O6HF46CFJNXIFM95X1HY/VCNP4190.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841331290-BF571JL4NBC4HE2X0AJ7/VCNP4191.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841349420-N0MFD2HYKCQBDRM04BNK/VCNP4215_1.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841406390-JK4V6AU01IPXI3WGP4H4/VCNP4216.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841367368-GX1SQBF21PAR27KY5HIM/VCNP4217.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841385269-PUDK66S4G495F49ODBHL/VCNP4218_1.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841402633-38T4O53OFXUY6JBUDCZE/VCNP4220.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841420481-E3ZI47EYCT4MK0WI0BX7/VCNP4221_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841432246-1RVZAB8LCCF0D2WMQ3B3/VCNP4222.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841854598-5ALAT583UE90R0UG4X6L/VCNP4223.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841806655-1KVD73YRR6DZRCQ8CX2H/VCNP4224_1.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841828073-WGQXPQQJBUMW82FUDYZE/VCNP4237.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841846905-V5KPAZT31UA21ODTDZPK/VCNP4238.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841863883-OI2EH8NNPBHAXHLN759M/VCNP4240.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841914855-69BED31SEF5OI45RP51T/VCNP4241_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841880778-36F17AOMSJGQLICDI04I/VCNP4242.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841897955-IZS5N6FM924VMV97GQ1B/VCNP4243.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841918795-PPI277RRY9SLGH8XBMU3/VCNP4244.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841982456-48A0F2ACWN9AW62FHRSR/VCNP4261_1_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841937948-Q102NIKGLSJ4I2RZNZG7/VCNP4263.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841960221-X38IPBWTERLG2VTUG798/VCNP4267.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841978838-XCCMA1AZPTQM67Q7TBW9/VCNP4270.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507841998399-ZAV2CJE6NRJ5FNZVB815/VCNP4271.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842171562-Z2NWDF0WOQF2I2P3QFXR/VCNP4272.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842128832-UP97LZ5PKJ0L91WQSIBK/VCNP4303.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842148705-4J5HMCLRPWVY669PSDE9/VCNP4307_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842166040-2GABLS7JGQOIEM6AIUM3/VCNP4538_1_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842184653-U94NTDIYXKX3EA4XNC3C/VCNP4558_1_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842224172-2ZNE4G5EUNHYSMKUZG61/VCNP4884_1_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842205460-2H4Q4685QFD3Q6U9FUXP/VCNP4899_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842258991-KXL6JJV2VECBIG2VMAKL/VCNP4908_1_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842245880-7XYL9XH7QV8JAXB72BDO/VCNP4913_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842295904-TE57UEE55JJPDUSKOWP6/VCNP5007_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842282890-GMTG6MS31Z11K6EC4OGB/VCNP5012_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842344636-HPYJFTM8JUFIKT53Y775/VCNP5026_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842319128-1UGN4WXKBP4JU8TQ9CV7/VCNP5105_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842336937-L94YBA03I5K0AIBUBTV8/VCNP5241_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842426841-VXO7Y13WXDVGFYFVQTA8/VCNP5263_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842382758-P2WPW56HODE2ULYB9L48/VCNP5484_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842401532-MKCO9NLB23KQEKSFYMQT/VCNP5501_1_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507842423911-3F0U1JRGBTSEVREFXA19/VCNP5551_1_FAV.JPG OUR SUBJECT ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504887444039-O77KPQSPBLSQ588UNL3W/VCNP3042_1_FAV.JPG OUR SUBJECT A disagreement between mothers. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505764461355-6QC0S5PJ04IJRJ1180G5/VCNP2243_1_FAV.JPG OUR SUBJECT - CHILDCARE https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504379804821-LMU7FKP87VLQZSD85YY3/EMBond-3782.small.jpg OUR SUBJECT Communal nursing in Utah prairie dogs. Photo by Elaine Miller Bond (www.elainemillerbond.com) https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504379614216-UPCQ89O3PTUD7A6LWKDF/EMBond-7148.small.jpg OUR SUBJECT Utah prairie dog female Backstripe allows her own offspring as well as nondescendant offspring to nurse from her. Photo by Elaine Miller Bond (www.elainemillerbond.com) https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504114524346-6M82IBDL6WOR3XVEX6IE/VCNP9061_coyote.JPG OUR SUBJECT A coyote has been spotted by the prairie dogs it's been stalking. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504114535921-PA8AU6910HVE3MDP597F/VCNP5266_coyote.JPG OUR SUBJECT A coyote hunts in the taller grass. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116513677-KPRZF73TU4WGUBGRWQGW/IMG_1619.JPG OUR SUBJECT A long-tailed weasel can make quick and easy prey of juvenile prairie dogs. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116507678-EFOINRDQFSFJUYMUJ6U5/IMG_1550.JPG OUR SUBJECT A wandering weasel makes its way into a prairie dog trap. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504118692508-7PC3SL8MUO7E6FJ5884J/IMG_3051.JPG OUR SUBJECT Long-tailed weasels sport white coats in the winter - the better to come upon their prey by surprise. ©John Hoogland 2010 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116522228-816ZIB2GOLIQ3GAIBICH/IMG_1623.JPG OUR SUBJECT Weasels are inquisitive animals. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116548081-C0WSOXUZTN2H084K2CKT/IMG_1658.JPG OUR SUBJECT Prairie dogs will sometimes (though infrequently) give false alarms at animals that are not predators before the prairie dog can determine what the animal is. Here another prey animal, the pronghorn, shows its own vulnerability. ©John Hoogland 2006 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116530659-PQD47Z3PBJLEZTBN1QWU/IMG_1993_1.JPG OUR SUBJECT Badgers are brazen predators in prairie dog colonies. ©John Hoogland 2007 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116540312-IT3JMM1LC8WO1JVP4701/IMG_3875.JPG OUR SUBJECT A Swainson's hawk lands in a colony during its hunt. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116549960-XTBPTFKTRURMMWMS68QL/IMG_3934.JPG OUR SUBJECT A Swainson's hawk flies over the colony looking for juvenile prairie dogs. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116568888-B3XGUYID9MHPYWE8CXSA/IMG_4512.JPG OUR SUBJECT Living within the confines of a prairie dog colony gives predators like badgers easier access to meaty prey. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116560147-YMG8WOH0N88SIBWW41O9/IMG_4528.JPG OUR SUBJECT A badger comes home with a juvenile prairie dog in her jaws. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116592423-XK8P89SZQ34D16B64K11/IMG_4620.JPG OUR SUBJECT A badger comes home with an adult prairie dog in her jaws. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116585890-4ZAWA0MQ5OW9E7REQ8YV/IMG_4542.JPG OUR SUBJECT This juvenile bullsnake will grow to adulthood hunting juvenile prairie dogs. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116576271-LYUHF13H7ACQNOJP9Y8R/IMG_4591.JPG OUR SUBJECT A gull from a nearby lake scavenges on the leftovers of a killed prairie dog. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116596432-Q1PV132B8SY95B46I7BE/IMG_4770.JPG OUR SUBJECT This badger mother must hunt almost nonstop to feed her growing offspring. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116636408-34OIZJ8QGQI3QEBY09WS/IMG_4827_1.JPG OUR SUBJECT A yawn puts this badger's crushing teeth on display. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116607646-6DKZC17Q4H794I7MH53W/IMG_5289_2.JPG OUR SUBJECT As a carnivore, badgers have long, sharp canines for grabbing prey. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116613901-GMYA7ELM3412ALHFQGHR/IMG_5759.JPG OUR SUBJECT This taxidermied stuffed badger is pulled through the colony to take targeted data on alarm calls. ©MRR 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116630651-N3LQR2LNWBJ82WTZ8AJS/VCNP2034.JPG OUR SUBJECT Coyotes are colored to blend in with their surroundings. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116658462-L2MPF6YOPR0I9G3Q8B0C/VCNP3021.JPG OUR SUBJECT Juvenile prairie dogs will often be startled by swooping songbirds like this mountain bluebird, being hypersensitive to their new surroundings. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116653024-OG917UVN0TZOPO0WZARW/VCNP5052.JPG OUR SUBJECT Coyotes are quick and silent, but find catching prairie dogs difficult because of colony vigilance. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116675694-MB6LTA6L12KTS2CF95MJ/VCNP5442.JPG OUR SUBJECT Ravens are more likely to scavenge on prairie dog carcasses, but will also opportunistically grab newly-emerged juvenile prairie dogs. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504116686423-YR2S8XHKM3BMRPS4KG5A/VCNP8291.JPG OUR SUBJECT A vulture soars overhead, searching for any carcasses leftover from predations. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505839881997-ACZRKI431S0MCABSD7BW/IMG_4919.JPG OUR SUBJECT - THE PREY ANIMAL https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504112767010-4DW1D41PG0LT0NBCFWVL/VCNP0484_alert.JPG OUR SUBJECT Upon perceiving a threat, a prairie dog will often run to the mouth of its burrow, keeping an eye on its surroundings from the safest position. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504112752852-X3Q5O1WEONI8LIWU9P6X/VCNP2446_FAV_alert.JPG OUR SUBJECT A juvenile prairie dog practices standing on alert. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504112857985-4EU27K2553NII1O9WNK9/GOPR0126_alert.JPG OUR SUBJECT Alert from the burrow mound. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504112786249-Y8J6SDNWMY7V9OLZ0WK6/VCNP9568_alert.JPG OUR SUBJECT Two yearling males stand in twin postures on alert. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504112711926-UOARB07A7XJTRYLSWKQT/VCNP3005_FAV_alert.JPG OUR SUBJECT Responding to an alarm call in the clan, almost every member stands on alert to locate the predator. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504112837264-WF6PLC00MFPKKHAJOJUA/VCNP5165_alert.JPG OUR SUBJECT Male 0 (zero) stands as tall as he can on alert. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504725881719-MRX8DSOA3EQXCLIQNEFL/IMG_5712_FAV.JPG OUR SUBJECT Three lactating mothers on alert. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504112848545-1B0NMVVKH53IDLAF7VXB/VCNP1888_alert.JPG OUR SUBJECT Standing at its tallest allows a prairie dog to scan farther. ©MRR 2017 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504725959705-KC0GTX0Y8HBQHBB9556L/IMG_4189_FAV.JPG OUR SUBJECT An entire clan on alert. ©John Hoogland 2011 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507134915885-KQ41K90JMD6LFTCITPKI/IMG_0074.JPG OUR SUBJECT Our menagerie of stuffed predators: a coyote, a bobcat, and two badgers. ©John Hoogland 2016 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504725529683-XU2D1LT67HLIDI02EBJU/IMG_4997_1_FAV.JPG OUR SUBJECT A badger decoy is pulled through the colony to measure alarm call responses. ©John Hoogland 2012 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493157849696-I1I0MCKR6XBG36N0C29H/VCNP2338_FAV.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1493940733087-HMGCR01YI2V95UPTNKOP/VCNP5578_FAV.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1495746655110-GU0EZN7RS7HVHLY70E82/IMG_4640_FAV.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504187945750-MA000QIDLNC2AYBRSNIX/IMG_4919.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504647329406-5K5RZ5GD0R044J8MRAMK/IMG_4893.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504731618247-HIOPR427LIBB8AE11JWL/IMG_4299_FAV.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505680227065-PKB3J9T4264159FS6TVY/VCNP2243_1_FAV.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505750355720-XU0ML256DU060AODBMO3/VCNP6554_1_FAV.JPG OUR SUBJECT https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1507844404000-HEVHFST76OF1TNIO3R7V/VCNP8417.JPG OUR SUBJECT https://www.prairiedoghoogland.com/videos-index daily 0.75 2017-10-04 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1505766140672-0O5CRYIZ4XUKFZCKRX9S/1219621.jpg VIDEOS - 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Title ©MRR 2017 https://www.prairiedoghoogland.com/videos daily 0.75 2019-01-01 https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504203098334-FZKOLMR8BJHASZC8OZZM/IMG_7823.JPG VIDEO GALLERIES https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504203746002-JPX0JI4LGP9MBIEXODHB/VCNP6167.JPG VIDEO GALLERIES https://images.squarespace-cdn.com/content/v1/5841a4e029687f506e15343d/1504224955203-7JTE5NAW7RYIDQ6JV9JR/image-asset.jpeg VIDEO GALLERIES - At a Nursery Burrow Adult male 31 inspects the GoPro while female baby 2x explores her surroundings. 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